Morphometry of Striped Tree Frogs, Polypedates leucomystax (Gravenhorst, 1829) from Indonesia with Description of a New Species

Awal Riyanto, Mumpuni, Jimmy A. McGuire


We examined patterns of geographic variation within Polypedates leucomystax using multivariate statistical analyzes of 14 morphological characters measured from 146 specimens obtained from Sumatra, Java, Kalimantan, and Sulawesi. Discriminant analysis revealed ten morphological characters and found that the Sulawesi population is morphologically distinct from all others considered. Following the Evolutionary Species Concept, we describe the Sulawesi population as a new species in P. leucomystax species complex.


geographic variation; Polypedates leucomystax; multivariate; Sulawesi; new species

Full Text:



Devitt T. J., LaDuc T. J., and McGuire J. A. (2008), «The Trimorphdon biscutatus (Squamata: Colubridae) species complex revisited: a multivariate statistical analysis of geographic variation», Copeia, 2008(2), 370 – 387.

Frost D. R. (ed.) (1985), Amphibian Species of the World. A Taxonomic and Geographic References, Allen Press, Kansas.

Frost D. R., Grant T., Faivovich J., Bain R. H., Haas A., Haddad C. F. B., de Sá R. O., Channing A., Wilkinson M., Donnellan S. C., Raxworthy C. J., Campbell L. A., Blotto B. L., Moler P., Drewes R. C., Nussbaum R. A., Lynch J. D., Green D. M., and Wheeler W. C. (2006), «The amphibian tree of life», Bull. Am. Mus. Nat. Hist., 297, 1 – 370.

Inger R. F., Stuart B. L. and Iskandar D. T. (2009), «Systematics of widespread Southeast Asian frog, Rana chalconota (Amphibian: Anura: Ranidae)», Zool. J. Linn. Soc., 155, 123 – 147.

Inger R. F. (1966), «The systematic and zoogeography of the Amphibians of Borneo», Fieldiana Zool. New. Ser., 52, 1 – 402.

Inger R. F. and Stuebing R. B. (2005), Field Guide to the Frogs of Borneo. Vol. II, Natural History Publisher (Borneo), Kinabalu.

Iskandar D. T. (1998), The Amphibians of Java and Bali, Research and Development Centre for Biology – LIPI – GEF – Biodiversity Collection Project, Bogor.

Iskandar D. T. and Colijn E. (2000), «Preliminary checklist of Southeast Asian and New Guinean herpetofauna. I. Amphibians», Treubia, 31(3), 1 – 133.

Kamsi M. (2003), Panduan Lapangan Amfibi Kawasan Ecosystem Leuser, The Gibbon Foundation and PILI – NGO Movement, Bogor.

Koch A., Arida E., Schmitz A., Böhme W., and Zigler T. (2009), «Refining the polytypic species concept of mangrove monitors (Squamata: Varanus indicus group): a new cryptic species from the Talaud Island, Indonesia, reveals the underestimated diversity of Indo-Australian monitor lizards», Austral. J. Zool., 57, 29 – 40.

Liem D. S. S. (1971), «The frogs and toads of Tjibodas National Park, Mt. Gede, Java, Indonesia», Phil. J. Sci., 100(4), 131 – 160.

McGarigal K., Cushman S., and Stafford S. G. (2000), Multivariate Statistics for Wildlife and Ecology Research, Springer-Verlag, New York.

Marquez R. and Eekhout X. R. (2006), «Advertisement calls of six species of anurans from Bali, Republic of Indonesia», J. Nat. Hist., 40(9 – 10), 571 – 588.

Menzies J. I. (1987), «A taxonomic revision of Papuan Rana (Amphibia: Ranidae)», Austral. J. Zool., 35, 373 – 418.

van Kampen P. N. (1923), The Amphibians of the Indo-Australian Archipelago, J. Brill, Leiden.

van Rooijen and Vogel G. (2008), «Contributions to a review of the Dendrelaphis pictus complex (Serpentes: Colubridae). 1. Description of a sympatric species», Amphibia– Reptilia, 29, 101 – 115.



  • There are currently no refbacks.

You can subscribe to the print or electronic version of the journal on the site of EastView Company. If you have any questions, please write to the email