Walterinnesia aegyptia Lataste, 1887 (Ophidia: Elapidae) and the Status of Naja morgani Mocquard 1905

Göran Nilson, Nasrullah Rastegar-Pouyani


We separate the eastern populations of Walterinnesia in Iran, Iraq and eastern Saudi Arabia under the name Walterinnesia morgani. This taxon is easily separated from Walterinnesia aegyptia in Egypt, Israel, western Saudi Arabia, and Jordan by a significantly lower number of anterior dorsal scale rows. Walterinnesia morgani is also having on average a lower number of ventrals, subcaudals, and united subcaudals, as well as a completely different juvenile color pattern, consisting of 25 to 33 reddish crossbars on an otherwise black body. In Walterinnesia aegyptia the juvenile dorsal color is black without any pattern.


Reptilia; Serpentes Elapidae; Bungarinae; Walterinnesia aegyptia; Walterinnesia morgani; taxonomy; Iran

Full Text:



Amr Z. S. and Amr S. S. (1983), «Snakebites in Jordan», Snake, 15, 81 – 85.

Anderson J. (1898), Zoology of Egypt. Vol. 1. Reptilia and Batrachia, London.

Anderson S. C. (1963), «Amphibians and reptiles from Iran», Proc. California Acad. Sci. Ser. 4, 31, 417 – 498.

Bogert C. M. (1943), «Dentitional phenomena in cobras and other elapids with notes on adaptive modification of fangs», Bull. Am. Mus. Nat. Hist., 81, 285 – 360.

Corcill N. L. (1932), «The snakes of Iraq», J. Bombay Nat. Hist. Soc., 35, 550 – 572.

Disi A. M., Modrý D., Necras P., and Rafai L. (2001), Amphibians and Reptiles of the Hashemite Kingdom of Jordan, Edition Chimaira.

Flower S. S. (1923), «On additions to the snake fauna of Egypt», Proc. Zool. Soc. Lond., 101, 1079–1083.

Flower S. S. (1933), «Notes on the recent reptiles and amphibians of Egypt, with a list of the species recorded from that kingdom», Proc. Zool. Soc. Lond., 111, 735–851.

Gasperetti J. (1988), «Snakes of Arabia», in: Fauna Saudi Arabia, Vol. 9, pp. 169 – 450.

Haas G. and Werner Y. L. (1969), «Lizards and snakes from southwestern Asia», Bull. Mus. Comp. Zool., 139(6), 327 – 406.

Joger U. (1984), «The venomous snakes of the Near and Middle East», in: Beihefte zum Tübinger Atlas des Vorderen Orients. Reihe A, No. 12.

Leviton A. E., Anderson S. C., Adler K., and Minton S. A. (1992), Handbook to Middle East Amphibians and Reptiles. Vol. 8, SSAR, Athens (OH, USA).

Marx H. (1953), «The elapid genus of snakes Walterinnesia», Fieldiana Zool., 34(16), 189 – 196.

Mertens R. (1929), «On rare snakes of the Senckenberg Museum», Bull. Antiv. Inst. Am., 3, 41.

Reed C. A. and Marx H. (1959), «A herpetological collection from northeastern Iraq», Trans. Kansas Acad. Sci., 62, 91 – 122.

Uğurtaş I. H., Papenfuss T. J., and Orlov N. L. (2001), New record of Walterinnesia aegyptia Lataste, 1887 (Ophidia: Elapidae: Bungarinae) in Turkey», Russ. J. Herpetol., 8(3), 239 – 245.



  • There are currently no refbacks.

You can subscribe to the print or electronic version of the journal on the site of EastView Company. If you have any questions, please write to the email